The independent role of morphology in mate choice is revealed by the rare instances where the usual association between song and morphology is disrupted.
The divergence of songs in the new population away from those in the progenitor population would only be prevented if these processes were balanced by repeated immigration and subsequent breeding: song flow.
To summarize, the particular song a male sings, and the behavioral responses of females to song and morphological signals, are not genetically inherited in a fixed manner but are determined by learning early in life.
Almost nothing is known from hybridization studies about the inheritance of courtship behavior of females, or of their responsiveness to particular male signals.
Thus the genetic basis to the origin of bird species is to be sought in the inheritance of adult traits that are subject to natural and sexual selection.